Entomology

Subject abbreviation: ENTM College of Natural and Agricultural Sciences

Christiane Weirauch, Ph.D., Chair Erin Wilson Rankin, Ph.D., Vice Chair Department Office, 175 Entomology insects.ucr.edu

Graduate Student Affairs

• (800) 735-0717 or (951) 827-1106

insects.ucr.edu/graduate-studies-program

Undergraduate Academic Advisor

• (951) 827-4186

insects.ucr.edu/undergrad-programs

Professors Michael E. Adams, Ph.D., Distinguished Professor, Entomology/Molecular, Cell and Systems Biology Peter W. Atkinson, Ph.D. Boris Baer, Ph.D. Dong-Hwan Choe, Ph.D. Brian A. Federici, Ph.D., Distinguished Professor of Graduate Division Alec Gerry, Ph.D. John M. Heraty, Ph.D. Chow-Yang Lee, Ph.D., Endowed Chair in Urban Entomology Quinn McFrederick, Ph.D. Jessica Purcell, Ph.D. Richard A. Redak, Ph.D. Michael K. Rust, Ph.D., Distinguished Professor of Graduate Division John T. Trumble, Ph.D., Distinguished Professor of Graduate Division Christiane Weirauch, Ph.D. Erin Wilson Rankin, Ph.D. Naoki Yamanaka, Ph.D.

Professors Emeriti Thomas S. Bellows, Jr., Ph.D. Ring T. Cardé, Ph.D. Elizabeth Grafton-Cardwell, Ph.D. J. Daniel Hare, Ph.D. E. Fred Legner, Ph.D. Jocelyn G. Millar, Ph.D. Thomas A. Miller, Ph.D. Joseph G. Morse, Ph.D. Bradley Mullens, Ph.D. Timothy D. Paine, Ph.D. Thomas M. Perring, Ph.D. John D. Pinto, Ph.D. Alexander Raikhel, Ph.D. Richard Stouthamer, Ph.D. S. Nelson Thompson, Ph.D. P. Kirk Visscher, Ph.D. Gregory P. Walker, Ph.D.

Associate Professors Allison Hansen, Ph.D. Kerry Mauck, Ph.D., Alfred M. Boyce Chair in Entomology Hollis Woodard, Ph.D. Assistant Professors Karthikeyan Chandrasegaran, Ph.D. Luciano Cosme, Ph.D. Ysabel Giraldo, Ph.D. Zachary MacDonald, Ph.D. Amy Murillo, Ph.D. Ikju Park, Ph.D. Elizabeth Rowen, Ph.D.

Morgan Thompson, Ph.D.

**

Professors of Cooperative Extension Bodil, Cass, Ph.D. Matthew Daugherty, Ph.D. Mark Hoddle, Ph.D., Distinguished Professor of Cooperative Extension S. Houston Wilson, Ph.D.

Cooperating Faculty Kurt Anderson, Ph.D., Evolution, Ecology, and Organismal Biology Alan Brelsford, Ph.D., Evolution, Ecology, and Organismal Biology Marko J. Spasojevic, Ph.D. Evolution, Ecology and Organismal Biology Linda Walling, Ph.D, Botany and Plant Sciences

The Department of Entomology offers under- graduate programs leading to either the B.S. or the B.A. degree. The B.S. degree offers students with a strong interest in the natural sciences an opportunity to emphasize this aspect of their education. The B.A. degree is available to stu- dents who wish to obtain a broader background in the humanities and social sciences than is required of students in the B.S. program.

The department offers six concentrations to choose from: General Insect Biology, Medi- cal, Public Health and Epidemiology, Insect Molecular Biology and Genomics, Conservation, Ecology and Evolution, Applied Entomology/ Pest Management, and Science Education. All students majoring in Entomology are normally required to take the core curriculum.

Information on the programs and course requirements is available at CNAS Academic Advising Center, 1223 Pierce Hall. Counseling, course recommendations, and information on education and career goals are provided by the Undergraduate Faculty Advisors, Dr. Allison Hansen, 224 Entomology and Dr. John Heraty, 138 Entomology.

Transfer Selection Criteria Applicants to majors in the College of Natural and Agricultural Sciences are selected on the basis of academic preparation, as assessed by their GPA and the strength of preparation for the intended major. A GPA of at least 2.70 is required. (This is a baseline GPA for consideration and not a guarantee of admission.) The GPA cutoff for Transfer admissions will be set each year by the appropriate Divisional Dean in consultation with the Executive Committee and the Chairs, and may differ by program depending on Transfer enrollment capacity.

In addition, applicants will need to complete college courses comparable to (articulated to) the following list of UCR courses to meet selection criteria for this major. Courses must be completed with "C" grades or better.

• 1. BIOL 005A, BIOL 05LA or BIOL 020, BIOL 005B, BIOL 005C

• 2. CHEM 001A, CHEM 01LA, CHEM 001B, CHEM 01LB, CHEM 001C, CHEM 01LC

• 3. MATH 005B or MATH 007A or MATH 009A

Final placement into required calculus classes will depend on class articulations.

To ensure graduation within four years, potential applicants are also encouraged to complete college courses comparable to (articulated to) the following UCR courses. These courses are not required to transfer into the major but are requirements for the major. Courses must be completed with "C" grades or better.

• 1. STAT 010

• 2. PHYS 002A, PHYS 02LA, PHYS 002B, PHYS 02LB, PHYS 002C, PHYS 02LC (must be calcu-

lus-based)

• 3. CHEM 008A CHEM 08LA, CHEM 008B, CHEM 08LB, CHEM 008C, CHEM 08LC

• 4. MATH 005C or MATH 007B or MATH 009B

Students should visit assist.org for updat- ed and comprehensive major preparation requirements.

• 3. Organic chemistry (one-year lower-

division), each course completed with a grade of “C” or better

• 4. PHYS 002A, PHYS 02LA, PHYS 002B, PHYS 02LB PHYS 002C, and PHYS 02LC

• 5. PHYS 040A, PHYS 040B, and PHYS 040C

• 6. MATH 009C, MATH 010A, MATH 010B, and

MATH 046

Courses must be completed with a letter grade, with no grade lower than a “C.”

Students should visit assist.org for updated and comprehensive major preparation requirements.

See Undergraduate Studies section.

See College of Natural and Agricultural Sciences, Colleges and Programs section.

Some of the following requirements for the major may also fulfill some of the college’s breadth requirements. Consult with a department advisor for course planning.

The major requirements for both the B.A. and the B.S. degrees in Entomology are as follows:

• 1. Lower-division core requirements (40

units)

• a) BIOL 005A, BIOL 05LA or BIOL 020, BIOL 005B, BIOL 005C

• b) MATH 005B or MATH 007A or MATH 009A or MATH 09HA

• c) MATH 005C or MATH 007B or MATH 009B

or MATH 09HB

• d) CHEM 001A, CHEM 001B, CHEM 001C,

CHEM 01LA, CHEM 01LB, CHEM 01LC

• e) STAT 010

• 2. Upper-division core requirements (18

units)

• a) ENTM 100/BIOL 100, ENTM 107, ENTM 180, and 4 units in any combination of

ENTM 190, ENTM 197, ENTM 199, or ENTM 199H

• b) BIOL 102

General Insect Biology Concentration

• 1. Lower-division requirements (23 units)

• a) CHEM 008A or CHEM 08HA, CHEM 08LA

or CHEM 08HLA, CHEM 008B or CHEM 08HB, CHEM 08LB or CHEM 08HLB

• b) PHYS 002A or PHYS 02HA, PHYS 02LA or

PHYS 02HLA, PHYS 002B or PHYS 02HB, PHYS 002C or PHYS 02HC, PHYS 02LC or PHYS 02HLC

• 2. Upper-division requirements (28 units)

• a) ENTM 173/BIOL 173

• b) BIOL 107A

• c) BCH 100 or BCH 100H

• d) Sixteen (16) additional units of ento-

mology electives, which may include up to 2 additional units of ENTM 190, ENTM 197, ENTM 199 or ENTM 199H.

Medical, Public Health & Epidemiology Con- centration

• 1. Lower-division requirements (23 units)

• a) CHEM 008A or CHEM 08HA, CHEM 08LA

or CHEM 08HLA, CHEM 008B or CHEM 08HB, CHEM 08LB or CHEM 08HLB

• b) PHYS 002A or PHYS 02HA, PHYS 02LA

or PHYS 02HLA, PHYS 002B or PHYS 02HB, PHYS 02LB or PHYS 02HLB, PHYS 002C or PHYS 02HC, PHYS 02LC or PHYS 02HLC

• 2. Upper-division requirements (28 units)

• a) ENTM 173/BIOL 173

• b) BIOL 107A

• c) BCH 100 or BCH 100H

• d) Sixteen (16) additional units of En-

tomology electives from the following courses: ENTM 101, ENTM 111, ENTM 114, ENTM 123, ENTM 125/ENTX 125/PLPA 125, ENTM 126, ENTM 133, ENTM 154/BIOL 154, BIOL 128, BIOL 157, BIOL 158: which may include up to 2 additional units of ENTM 190, ENTM 197, ENTM 199 or ENTM 199H.

Insect Molecular Biology & Genomics Concen- tration

• 1. Lower-division requirements (23 units)

• a) CHEM 008A or CHEM 08HA, CHEM 08LA

or CHEM 08HLA, CHEM 008B or CHEM 08HB, CHEM 08LB or CHEM 08HLB

• b) PHYS 002A or PHYS 02HA, PHYS 02LA

or PHYS 02HLA, PHYS 002B or PHYS 02HB, PHYS 02LB or PHYS 02HLB, PHYS 002C or PHYS 02HC, PHYS 02LC or PHYS 02HLC

• 2. Upper-division requirements (28 units)

• a) ENTM 173/BIOL 173

• b) BIOL 107A

• c) BCH 100 or BCH 100H

• d) Sixteen (16) additional units of En-

tomology electives from the following courses: ENTM 101, ENTM 106, ENTM 111, ENTM 112/BIOL 112/BPSC 112, ENTM 123, ENTM 125/ENTX 125/PLPA 125, ENTM 126, BIOL 118, BIOL 121, BIOL 128, GNBT 110, GNBT 114; which may include up to two (2) additional units of ENTM 190, ENTM 197, ENTM 199 or ENTM 199H.

Conservation, Ecology & Evolution Concen- tration

• 1. Lower-division requirements (17-18 units)

• a) CHEM 008A or CHEM 08HA, CHEM 08LA

or CHEM 08HLA, CHEM 008B or CHEM 08HB, CHEM 08LB or CHEM 08HLB

• b) PHYS 002A or PHYS 02HA, PHYS 02LA

or PHYS 02HLA, and either PHYS 002B or PHYS 02HB, and PHYS 02LB OR PHYS 02HLB; or PHYS 008

• 2. Upper-division requirements (24 units)

• a) ENTM 173/BIOL 173

• b) BCH 100 or BCH 100H

• c) Sixteen (16) additional units of Ento-

mology electives from the following courses: ENTM 101, ENTM 106, ENTM 108, ENTM 109, ENTM 112/BIO 112/BPSC 112, ENTM 114, ENTM 123, ENTM 127/BIOL 127, ENTM 130, ENTM 133, ENTM 139/MCBL 139, ENTM 162/BIOL 162, BIOL 151, BIOL 166; which may include up to two (2) additional units of ENTM 190, entm 197, ENTM 199 OR ENTM 199H.

Applied Entomology/Pest Management Con- centration

• 1. Lower-division requirements (17-18 units)

• a) CHEM 008A or CHEM 08HA, CHEM 08LA

or CHEM 08HLA, CHEM 008B or CHEM 08HB, CHEM 08LB or CHEM 08HLB

• b) PHYS 002A or PHYS 02HA, PHYS 02LA

or PHYS 02HLA, and either PHYS 002B or PHYS 02HB, and PHYS 02LB or PHYS 02HLB; or PHYS 008

• 2. Upper-division requirements (24 units)

• a) ENTM 173/BIOL 173

• b) BIOL 100 or BCH 100H

• c) Sixteen (16) additional units of Ento-

mology electives from the following courses: ENTM 106, ENTM 112/BIOL 112/ BPSC 112, ENTM 114, ENTM 123, ENTM 124, ENTM 125/ENTX 125/PLPA 125, ENTM 126, ENTM 128, ENTM 129, ENTM 130, ENTM 133; which may include up to two (2) additional units of ENTM 190, ENTM 197, ENTM 199 or ENTM 199H.

Science Education Concentration Students must consult with their Entomology advisor before electing this option.

• 1. Lower-division requirements (4-5 units)

• a) One of PHYS 008 or PHYS 010 or (PHYS 002A or PHYS 02HA, PHYS 02LA or PHYS

02HLA)

• 2. Upper-division requirements (32 units)

• a) EDUC 132, EDUC 147, EDUC 178, EDUC 179A

• b) Sixteen (16) additional units of Ento-

mology electives, which may include up to 2 additional units of ENTM 190, ENTM 197, ENTM 199 or ENTM 199H.

For lower-division students interested in pur- suing the Science Education Concentration in Entomology, EDUC 001 is recommended.

Upper division courses in BIOL, BPSC, and related programs including but not limited to BIOL 151 and BPSC 133 are suggested to acquire a background in the life sciences appropriate for an Entomology major.

For students intending to specialize at the graduate level in insect toxicology or insect physiology, biochemistry, and molecular biology, it is recommended that BCH 110A, BCH 110B, and BCH 110C sequence and BCH 102 be substituted in place of an equal number of upper-division course units in life sciences. Due to course context overlap, credit is not awarded for BCH 110A, BCH 110B, or BCH 110C if it has already been awarded for BCH 100

The Department of Entomology offers a minor in Entomology designed to allow the student the freedom to pursue areas of particular interest.

The minor consists of no less than 20 and no more than 28 units of Entomology courses to be selected as follows:

• 1. ENTM 100/BIOL 100

• 2. Select from the following upper-division

Entomology courses to complete unit requirement: ENTM 106, ENTM 107, ENTM 109, ENTM 112/BIOL 112/BPSC 112, ENTM 114, ENTM 124, ENTM 125, ENTM 126, ENTM 127/BIOL 127, ENTM 129, ENTM 129L, ENTM 133, ENTM 154, ENTM 162/BIOL 162, ENTM 173/BIOL 173, ENTM 180, ENTM 190, ENTM 197, ENTM 199, ENTM 199H

• 3. No more than 4 units of ENTM 190, ENTM 197, ENTM 199, or ENTM 199H, either solely

or in combination, may be applied toward the unit requirement.

• 4. Of the specified upper-division units, a

minimum of 16 must be unique to the minor and may not be used to satisfy major requirements.

See Minors under the College of Natural and Agricultural Sciences in the Colleges and Programs section of this catalog for additional information on minors.

The College of Natural and Agricultural Science offers a combined five-year B.S.+M.S. program in Entomology, designed to allow successful UCR Entomology B.S. graduates to complete the Master of Science degree in Entomology in one year (Plan I, Thesis).

Applicants to the Combined Entomology B.S.+M.S. program (including transfer students) should apply by the end of their junior year, providing that the student is a UCR Entomolo- gy B.S. student with cumulative GPA at least 3.0 overall and 3.3 GPA in the Entomology major. The application to the Combined B.S.+M.S. pro- gram must include a Statement of Interest and Eligibility, at least two recommendation letters from UCR Academic Senate faculty members, one of which should include the prospective thesis advisor and at least one of which must be Entomology faculty. Matriculation into the graduate portion of the combined degree pro- gram occurs in the Fall term following senior year, provided: (a) the M.S. application is ac- cepted, (b) throughout senior year, the student is an Entomology B.S. major with cumulative GPA 3.0 or higher, (c) by the end of senior year, the student completes the Entomology B.S. degree requirements.

Incoming freshman students who apply to the Entomology B.S. program may simulta- neously apply for preliminary admission into the combined degree program provided their high-school GPA is at least 3.6, they satisfy the Entry-Level Writing requirement prior to matriculation, and they have sufficient math preparation to enroll in MATH 7A (Calculus for the Life Sciences) or 9A (First-Year Calculus Part 1) upon arrival. Preliminary conditional admission status is maintained as long as the student is an Entomology B.S. student in good standing with a cumulative GPA of at least 3.0. Conditionally admitted students still need to apply for full admission by the end of their junior year as described above.

Overall Requirements A total of 36 units are required to complete the MS portion. 24 units must be 200-level. No more than 12 units may come from 297 or 299 research units. No more than 12 units of ENTM 100-level courses earned prior to matricula- tion to graduate status can be applied toward the MS degree requirements.

Course Requirements

• 1. While in the Combined Entomology

B.S.+M.S. Program, students must com- plete a minimum of 24 units of research (a combination of ENTM 190/197/199 and ENTM 297/299) over 6 consecutive quarters. Students receive credit towards this requirement by completing up to 12 units of ENTM 190/197/199/199H as an undergraduate student and a minimum of 12 units of ENTM 297/299 as a graduate student. Only 12 units of ENTM 297/299 will count toward the 36 unit requirement.

• 2. A minimum of 9 units of graduate courses

and/or graded 2-unit graduate seminars from this list of courses: ENTM 201, ENTM 201L, ENTM 202, ENTM 202L, ENTM 203, ENTM 203L, ENTM 210, ENTM 211, ENTM 212, ENTM 219, ENTM 227, ENTM 229, ENTM 230, ENTM 240, ENTM 241, ENTM 242, ENTM 249, ENTM 251, ENTM 252, ENTM 253, ENTM 254, ENTM 255, ENTM 256, ENTM 257, ENTM 258, ENTM 259, ENTM 260, ENTM 262, ENTM 271, ENTM 272, ENTM 276, ENTM 277, ENTM 289, ENTM 290, ENTM 423, BPSC 230, BPSC 234, BPSC 246, BPSC 247, EEOB 215, EEOB 217, EEOB 230, EEOB 282, EEOB 283, STAT 231A, STAT 231B.

• 3. Up to 6 units of upper division 100 level

ENTM courses may be taken during the M.S. portion of the program.

• 4. Enrollment in ENTM 250, is required during

all quarters of M.S. study (3 units total).

• 5. Thesis and Final Oral Examination: Follow-

ing completion of their research, students submit a written thesis and conclude their studies with an oral public defense of the thesis. Interested students should check with the Department of Entomology's Undergraduate Advisors and their Academic Advisor for addi- tional details.

Professional Development Students in the Entomology B.S.+M.S. Program must participate in the departmental seminar (ENTM 250) for the three quarters of their mas- ter's year and present at the Annual Graduate Student Seminar Day.

The Department of Entomology offers programs leading to the M.S. (thesis plan) and Ph.D. degrees with specialization in, but not restricted to, the following areas of study:

• Arthropod vectors of human, animal, and plant pathogens

• Behavior

• Biochemistry and Physiology

• Chemical Ecology

• Conservation Biology and Global Change

• Endocrinology and Development

• Ecology and Evolution

• Genetics, Genomics, and Molecular Biology

• Insect Pathology

• Integrated Pest Management

• Invasive Species and Biological Control

• Medical and Veterinary Entomology

• Nematology

• Neuroscience

• Plant-Herbivore Interactions

• Social Insects and Pollination Biology

• Systematics

• Urban Entomology

Information on participating faculty and their research specializations may be found at insects.ucr.edu. University requirements for the M.S. and Ph.D. degrees are given in the Graduate Studies section of this catalog.

Admission For admission to the graduate program, prospective applicants must have a bachelor's degree with a major in Entomology, a biological science, Chemistry, Biochemistry, or a suitable equivalent. Regardless of undergraduate major, students must have strength in life sciences. Recommended (but not required for admission) courses vary depending on research focus and include biochemistry, chemistry and organic chemistry, data sciences, entomology, genetics, physics, statistics and other courses specific to research areas.

Credit from these courses does not count toward the unit requirement of the M.S. degree.

All applicants whose first language is not English and do not have an undergraduate or graduate degree from an accredited institution where English is the exclusive language of instruction are required to provide a valid and passing English Proficiency exam score. Overall minimum scores are: TOEFL: 80 (iBT); 550 for pBT. IELTS: overall score of 7.0, with no individual component score less than 6.0. Course Work All students must take ENTM 201, ENTM 201L, ENTM 202, ENTM 202L, ENTM 203, and ENTM 203L (15 units).

Seminar Requirements

• 1. All full-time students must enroll in the

ENTM 250 seminar during each quarter in which it is offered. (units vary)

• 2. Students in the second year and beyond

must present a seminar or a poster pre- sentation of their research at the Annual Graduate Student Seminar Day.

Professional Development Entomology students must complete an an- nual Individual Development Plan along with their annual progress report and present an annual seminar or poster presentation covering their research investigations at the Annual Graduate Student Seminar Day, and complete ENTM 201, ENTM 202, and ENTM 203.

Each student, with the advice of their Ph.D. Guidance Committee, will select courses that complement their research program and help the student prepare for the qualifying examination (units vary).

Students must take at least four Entomology 2-unit seminar courses for a letter grade from the following list: ENTM 249, ENTM 251, ENTM 252, ENTM 253, ENTM 254, ENTM 255, ENTM 256, ENTM 257, ENTM 258, ENTM 259, ENTM 260, ENTM 262, ENTM 289, ENTM 423 (8 units).

Written and Oral Qualifying Examinations Advancement to candidacy depends on the student passing written and oral qualifying examinations.

Dissertation and Final Oral Examinations Following completion of their research, stu- dents submit a written dissertation and con- clude their studies with an oral public defense of the dissertation.

Exam Modality In-person is the default modality for qualify- ing exams and final defenses. If unforeseen circumstances arise and the exam committee chair approves, a hybrid exam will be allowed. If a hybrid exam is approved, committee members not based at the UC Riverside campus may attend remotely but the majority of committee members must be physically present (e.g. 2 out of 3 for the final defense, 3 out of 5 for the qualifying exam). The exam committee chair and the student are required to be physically present.

Teaching Requirement Ph.D. students must fulfill a three quarter teaching requirement.

Normative Time to Ph.D. 17 quarters

Students should refer to the Graduate Studies section of the catalog for the minimum mas- ter’s degree requirements.

Each student, with the advice of their M.S. Guidance Committee, will select courses that complement their research program (units vary). Students must take at least two Entomology 2-unit seminar courses for a letter grade from the following list: ENTM 249, ENTM 251, ENTM 252, ENTM 253, ENTM 254, ENTM 255, ENTM 256, ENTM 257, ENTM 258, ENTM 259, ENTM 260, ENTM 262, ENTM 289, ENTM 423 (4 Units Total).

Thesis and Final Oral Examination Following completion of their research, students submit a written thesis and conclude their studies with an oral public defense of the thesis.

Normative Time to M.S. 6 quarters

Opportunities for Interdisciplinary Graduate Study Faculty from the Department of Entomology also participate in the following additional graduate programs:

• Biochemistry and Molecular Biology

• Cell, Molecular, and Developmental Biology (CMDB)

• Neuroscience

• Chemistry

• Environmental Toxicology

• Evolution, Ecology, and Organismal Biology (EEOB)

• Genetics, Genomics and Bioinformatics

These interdepartmental programs draw on the strengths of distinguished scientists from several units. For further information concern- ing work in these areas, see the respective program descriptions in the Programs and Courses section of this catalog or contact the CNAS Graduate Student Affairs Center, at (800) 735-0717.