Entomology

Subject abbreviation: ENTM College of Natural and Agricultural Sciences

Richard A. Redak, Ph.D., Chair Thomas M. Perring , Ph.D., Vice Chair Department Office, 175 Entomology insects.ucr.edu

Graduate Student Affairs

• (800) 735-0717 or (951) 827-5621

insects.ucr.edu/graduate-studies-program

Undergraduate Faculty Advisor

• (951) 827-5717

insects.ucr.edu/undergradute-programs- entomology Professors Michael E. Adams, Ph.D., Entomology/Molecular, Cell and Systems Biology Peter W. Atkinson, Ph.D. Boris Baer, Ph.D. Ring T. Cardé, Ph.D., Distinguished Professor, Alfred M. Boyce Chair in Entomology Brian A. Federici, Ph.D., Distinguished Professor of Graduate Division Alec Gerry, Ph.D. John M. Heraty, Ph.D. Chow-Yang Lee, Ph.D. Jocelyn G. Millar, Ph.D., Distinguished Professor Timothy D. Paine, Ph.D., Distinguished Professor, Tokuji and Bettie L. Furuta Endowed Chair Thomas M. Perring, Ph.D. Alexander Raikhel, Ph.D., Distinguished Professor, Mir Mulla Endowed Chair in Medical Entomology Richard A. Redak, Ph.D. Michael K. Rust, Ph.D., Distinguished Professor of Graduate Division Richard Stouthamer, Ph.D. John T. Trumble, Ph.D., Distinguished Professor of Graduate Division Christiane Weirauch, Ph.D.

Professors Emeriti Thomas S. Bellows, Jr., Ph.D. Leland R. Brown, Ph.D. Elizabeth Grafton-Cardwell, Ph.D. Richard D. Goeden, Ph.D. J. Daniel Hare, Ph.D. Marshall W. Johnson, Ph.D. E. Fred Legner, Ph.D. Thomas A. Miller, Ph.D. Joseph G. Morse, Ph.D. Mir S. Mulla, Ph.D. Bradley Mullens, Ph.D. John D. Pinto, Ph.D. S. Nelson Thompson, Ph.D. P. Kirk Visscher, Ph.D. Gregory P. Walker, Ph.D.

Associate Professors Allison Hansen, Ph.D. Dong-Hwan Choe, Ph.D. Quinn McFrederick, Ph.D. Jessica Purcell, Ph.D. Erin W. Rankin, Ph.D. Naoki Yamanaka, Ph.D.

Assistant Professors Ysabel Giraldo, Ph.D. Kerry Mauck, Ph.D. Amy Murillo, Ph.D. Hollis Woodard, Ph.D. **

Lecturers Matthew Daugherty, Ph.D., Integrative Biology Mark Hoddle, Ph.D., Biological Control S. Houston Wilson, Ph.D., Pest Management Monique Rivera, Ph.D. Pest Management

Cooperating Faculty Kurt Anderson, Ph.D., Evolution, Ecology, and Organismal Biology Alan Brelsford, Ph.D., Evolution, Ecology, and Organismal Biology Richard Cardullo, Ph.D., Evolution, Ecology, and Organismal Biology Anupama Dahanukar, Ph.D. Molecular, Cell and Systems Biology Paul De Ley, Ph.D., Nematology Paul Nabity, Ph.D., Botany and Plant Sciences Nicole Rafferty, Ph.D., Evolution, Ecology, and Organismal Biology Linda Walling, Ph.D, Botany and Plant Sciences

The Department of Entomology offers under- graduate programs leading to either the B.S. or the B.A. degree. The B.S. degree offers students with a strong interest in the natural sciences an opportunity to emphasize this aspect of their education. The B.A. degree is available to students who wish to obtain a broader background in the humanities and social sciences than is required of students in the B.S. program.

Information on the programs and course requirements is available at CNAS Academic Advising Center, 1223 Pierce Hall. Counseling, course recommendations, and information on education and career goals are provided by the Undergraduate Faculty Advisors, Dr. Dong-Hwan Choe, 382 Entomology and Dr. John Heraty, 138 Entomology.

Transfer Selection Criteria Applicants to majors in the College of Natural and Agricultural Sciences are selected on the basis of academic preparation, as assessed by their GPA and the strength of preparation for the intended major. A GPA of at least 2.70 is required. (This is a baseline GPA for consideration and not a guarantee of admission.) The GPA cutoff for Transfer admissions will be set each year by the appropriate Divisional Dean in consultation with the Executive Committee and the Chairs, and may differ by program depending on Transfer enrollment capacity.

In addition, applicants will need to complete college courses comparable to at least two of the following UCR year-long sequences in order to meet selection criteria for this major. Courses must be completed with “C” grades or better:

MATH 007A or MATH 009A and MATH 007B or MATH 009B (mandatory)

And at least one sequence from:

• 1. BIOL 005A, BIOL 05LA or BIOL 020 and

BIOL 005B (and BIOL 005C, if articulated)

• 2. CHEM 001A, CHEM 01LA, CHEM 001B, CHEM 01LB, CHEM 001C, and CHEM 01LC

• 3. Organic chemistry (one-year lower-

division), each course completed with a grade of “C” or better

• 4. PHYS 002A, PHYS 02LA, PHYS 002B, PHYS 02LB PHYS 002C, and PHYS 02LC

• 5. PHYS 040A, PHYS 040B, and PHYS 040C

• 6. MATH 009C, MATH 010A, MATH 010B, and

MATH 046

Courses must be completed with a letter grade, with no grade lower than a “C.” Students should visit assist.org for updated and comprehensive major preparation requirements.

See Undergraduate Studies section.

See College of Natural and Agricultural Scienc- es, Colleges and Programs section.

Some of the following requirements for the major may also fulfill some of the college’s breadth requirements. Consult with a depart- ment advisor for course planning.

The major requirements for both the B.A. and the B.S. degrees in Entomology are as follows:

• 1. Lower-division requirements (64 units)

• a) BIOL 005A, BIOL 05LA or BIOL 020, BIOL 005B, BIOL 005C

• b) PHYS 002A, PHYS 002B, PHYS 002C,

PHYS 02LA, PHYS 02LB, PHYS 02LC

• c) MATH 007A or MATH 009A, MATH 007B

or MATH 009B

• d) CHEM 001A, CHEM 001B, CHEM 001C,

CHEM 01LA, CHEM 01LB, CHEM 01LC, CHEM 008A or CHEM 08HA, CHEM 008B or CHEM 08HB, CHEM 008C or CHEM 08HC, CHEM 08LA or CHEM 8HLA, CHEM 08LB or CHEM 8HLB, CHEM 08LC or CHEM 8HLC.

• e) STAT 010

• 2. Upper-division requirements (46 units)

• a) ENTM 100/BIOL 100, ENTM 107, ENTM 173/BIOL 173, ENTM 180, and 4 units in

any combination of ENTM 190, ENTM 197, ENTM 199, or ENTM 199H

• b) Sixteen (16) additional units of ento-

mology electives, which may include up to 2 additional units of ENTM 190, ENTM 197, ENTM 199 or ENTM 199H

• c) BCH 100

• d) BIOL 102

• e) BIOL 107A

Upper division courses in BIOL, BPSC, and related programs including but not limited to BIOL 151 and BPSC 133 are suggested to acquire a background in the life sciences appropriate for an Entomology major.

For students intending to specialize at the graduate level in insect toxicology or insect physiology, biochemistry, and molecular biology, it is recommended that the BCH 110A, BCH 110B, and BCH 110C sequence and BCH 102 be substituted in place of an equal number of upper-division course units in life sciences. Due to course content overlap, credit is not awarded for BCH 110A, BCH 110B, or BCH 110C if it has already been awarded for BCH 100. Sample Program

Freshman Year Fall Winter Spring

BIOL 005A, BIOL 05LA or BIOL 020; BIOL 005B 4 4

CHEM 001A, CHEM 001B, CHEM 001C, CHEM 01LA, CHEM 01LB, CHEM 01LC 4,1 4,1 4,1

ENGL 001A, ENGL 001B, 4 4

MATH 007A or MATH 009A, MATH 007B or MATH 009B 4 4

Humanities/Social Sciences 4 4

Total Units 17 17 13

Sophomore Year Fall Winter Spring

BIOL 005C 4

Biology/Entomology Electives 4

CHEM 008A and 008LA, CHEM 008B and 008LB, CHEM 008C and 008LC 4 4 4

PHYS 02A, PHYS 02LA, PHYS 02B, PHYS 02LB, PHYS 02C, PHYS 02LC 4,1 4,1 4,1

Humanities/Social Sciences, STAT 010 4 4 5

Total Units 17 17 14

Junior Year Fall Winter Spring

BIOL 102 4

BIOL 107A, ENTM 173/ BIOL 173 4 4

ENTM 100/BIOL 100 4

ENTM 107 4

Biology/Entomology Electives 7 8

BCH 100, ENTM 19X 4 2

Humanities/Social Sciences 4

Total Units 16 13 16

Senior Year Fall Winter Spring

ENTM 180, ENTM 19X 2 2

Biology/Entomology Electives 8 8 8

Humanities/Social Sciences, ENGL 001C 4 4 4

Total Units 14 12 14

The Department of Entomology offers a minor in Entomology designed to allow the student the freedom to pursue areas of particular interest.

The minor consists of no less than 20 and no more than 28 units of Entomology courses to be selected as follows:

• 1. ENTM 100/BIOL 100

• 2. Select from the following upper-division

Entomology courses to complete unit requirement: ENTM 106, ENTM 107, ENTM 109, ENTM 112/BIOL 112/BPSC 112, ENTM 114, ENTM 124, ENTM 125, ENTM 126, ENTM 127/BIOL 127, ENTM 129, ENTM 129L, ENTM 133, ENTM 154, ENTM 162/BIOL 162, ENTM 173/BIOL 173, ENTM 180, ENTM 190, ENTM 197, ENTM 199, ENTM 199H

• 3. No more than 4 units of ENTM 190, ENTM 197, ENTM 199, or ENTM 199H, either solely

or in combination, may be applied toward the unit requirement.

• 4. Of the specified upper-division units, a

minimum of 16 must be unique to the minor and may not be used to satisfy major requirements.

See Minors under the College of Natural and Agricultural Sciences in the Colleges and Programs section of this catalog for additional information on minors.

The College of Natural and Agricultural Science offers a combined five-year B.S.+M.S. program in Entomology, designed to allow successful UCR Entomology B.S. graduates to complete the Master of Science degree in Entomology in one year (Plan I, Thesis).

Applicants to the Combined Entomology B.S.+M.S. program (including transfer students) should apply by the end of their junior year, providing that the student is a UCR Entomolo- gy B.S. student with cumulative GPA at least 3.0 overall and 3.3 GPA in the Entomology major. The application to the Combined B.S.+M.S. pro- gram must include a Statement of Interest and Eligibility, at least two recommendation letters from UCR Academic Senate faculty members, one of which should include the prospective thesis advisor and at least one of which must be Entomology faculty. Matriculation into the graduate portion of the combined degree pro- gram occurs in the Fall term following senior year, provided: (a) the M.S. application is ac- cepted, (b) throughout senior year, the student is an Entomology B.S. major with cumulative GPA 3.0 or higher, (c) by the end of senior year, the student completes the Entomology B.S. degree requirements.

Incoming freshman students who apply to the Entomology B.S. program may simulta- neously apply for preliminary admission into the combined degree program provided their high-school GPA is at least 3.6, their SAT-I combined score is at least 1250, they satisfy the Entry-Level Writing requirement prior to matriculation, and they have sufficient math preparation to enroll in MATH 7A (Calculus for the Life Sciences) or 9A (First-Year Calculus Part 1) upon arrival. Preliminary conditional admission status is maintained as long as the student is an Entomology B.S. student in good standing with a cumulative GPA of at least 3.0. Conditionally admitted students still need to apply for full admission by the end of their junior year as described above. Overall Requirements A total of 36 units are required to complete the MS portion. 24 units must be 200-level. No more than 12 units may come from 297 or 299 research units. No more than 12 units of ENTM 100-level courses earned prior to matricula- tion to graduate status can be applied toward the MS degree requirements.

Course Requirements

• 1. While in the Combined Entomology

B.S.+M.S. Program, students must com- plete a minimum of 24 units of research (a combination of ENTM 190/197/199 and ENTM 297/299) over 6 consecutive quarters. Students receive credit towards this requirement by completing up to 12 units of ENTM 190/197/199/199H as an undergraduate student and a minimum of 12 units of ENTM 297/299 as a graduate student. Only 12 units of ENTM 297/299 will count toward the 36 unit requirement.

• 2. A minimum of 9 units of graduate courses

and graded 2-unit graduate seminars from this list of courses: ENTM 201, ENTM 202, ENTM 203, ENTM 207, ENTM 209, ENTM 210, ENTM 212, ENTM 219, ENTM 227, ENTM 229, ENTM 230, ENTM 240, ENTM 241, ENTM 242, ENTM 249, ENTM 251, ENTM 252, ENTM 254, ENTM 255, ENTM 256, ENTM 257, ENTM 258, ENTM 262, ENTM 267, ENTM 289, ENTM 290, BPSC 230, BPSC 234, BPSC 246, BPSC 247, EEOB 215, EEOB 217, EEOB 230, EEOB 282, EEOB 283, STAT 231A, STAT 231B.

• 3. Up to 6 units of upper division 100 level

ENTM courses may be taken during the M.S. portion of the program.

• 4. Enrollment in ENTM 250, is required during

all quarters of M.S. study (3 units total).

• 5. Thesis and Final Oral Examination: Follow-

ing completion of their research, students submit a written thesis and conclude their studies with an oral public defense of the thesis.

Interested students should check with the Department of Entomology's Undergraduate Advisors and their Academic Advisor for addi- tional details.

Professional Development Students in the Entomology B.S.+M.S. Program must participate in the departmental seminar (ENTM 250) for the three quarters of their mas- ter's year and present at the Annual Graduate Student Seminar Day.

Graduate Program The Department of Entomology offers programs leading to the M.S. (thesis plan) and Ph.D. degrees with specialization in, but not restricted to, the following areas of study:

• Arthropod vectors of human, animal, and plant pathogens

• Behavior

• Biochemistry and Physiology

• Chemical Ecology

• Conservation Biology and Global Change

• Endocrinology and Development

• Ecology and Evolution

• Genetics, Genomics, and Molecular Biology • Insect Pathology

• Integrated Pest Management

• Invasive Species and Biological Control

• Medical and Veterinary Entomology

• Nematology

• Neuroscience

• Plant-Herbivore Interactions

• Social Insects and Pollination Biology

• Systematics

• Urban Entomology

Information on participating faculty and their research specializations may be found at insects.ucr.edu. University requirements for the M.S. and Ph.D. degrees are given in the Graduate Studies section of this catalog.

Admission Students must have a bachelor’s degree with a major in Entomology, a biological science, Chemistry, Biochemistry, or a suitable equiva- lent. Regardless of undergraduate major, stu- dents must have had, or complete soon after entering graduate school, the following:

• 1. One year of course work each in general

biology, general chemistry, and organic chemistry.

• 2. The equivalent of a one quarter course

each in genetics and biochemistry.

• 3. The equivalent of 30 quarter units of life

sciences other than entomology. Students who wish to specialize in insect biochem- istry, insect physiology, molecular ento- mology, neuroscience, or toxicology may substitute additional courses in physical, organic, and biological chemistry; toxicol- ogy; and pharmacology for courses in life sciences.

Credit from these courses does not count toward the unit requirement of the M.S. degree.

The department requires GRE General Test scores (verbal, quantitative, and analytical). All applicants whose first language is not English and do not have an undergraduate or graduate degree from an accredited institution where English is the exclusive language of instruction must submit a recent Test of English as a For- eign Language (TOEFL) and obtain a minimum score on the exam of 550 (paper-based), 213 (computer-based), or 80 (internet-based).

Course Work All students must take ENTM 201, ENTM 202, and ENTM 203 (14 units).

Seminar Requirements

• 1. All full-time students must enroll in the

ENTM 250 seminar during each quarter in which it is offered. (units vary)

• 2. Students in the second year and beyond

must present a seminar or a poster pre- sentation of their research at the Annual Graduate Student Seminar Day.

Professional Development Entomology students must complete an an- nual Individual Development Plan along with their annual progress report and present an annual seminar or poster presentation cover- ing their research investigations at the Annual Graduate Student Seminar Day, and complete ENTM 201.

Each student, with the advice of their Ph.D. Guidance Committee, will select courses that complement their research program and help the student prepare for the qualifying examination (units vary).

Students must take at least four Entomology 2-unit seminar courses for a letter grade from the following list: ENTM 249, ENTM 251, ENTM 252, ENTM 254, ENTM 255, ENTM 256, ENTM 257, ENTM 258, ENTM 262, ENTM 289 (8 units).

Written and Oral Qualifying Examinations Advancement to candidacy depends on the student passing written and oral qualifying examinations.

Dissertation and Final Oral Examinations Following completion of their research, stu- dents submit a written dissertation and con- clude their studies with an oral public defense of the dissertation.

Teaching Requirement Ph.D. students must fulfill a three quarter teaching requirement.

Normative Time to Ph.D. 17 quarters

Students should refer to the Graduate Studies section of the catalog for the minimum mas- ter’s degree requirements.

Each student, with the advice of their M.S. Guidance Committee, will select courses that complement their research program (units vary).

Students must take at least two Entomology 2-unit seminar courses for a letter grade from the following list: ENTM 249, ENTM 251, ENTM 252, ENTM 254, ENTM 255, ENTM 256, ENTM 257, ENTM 258, ENTM 262, ENTM 289 (4 Units).

Thesis and Final Oral Examination Following completion of their research, students submit a written thesis and conclude their studies with an oral public defense of the thesis.

Normative Time to M.S. 6 quarters

Opportunities for Interdisciplinary Graduate Study Faculty from the Department of Entomology also participate in the following additional graduate programs:

• Biochemistry and Molecular Biology

• Cell, Molecular, and Developmental Biology (CMDB)

• Neuroscience

• Chemistry

• Environmental Toxicology

• Evolution, Ecology, and Organismal Biology (EEOB)

• Genetics, Genomics and Bioinformatics

These interdepartmental programs draw on the strengths of distinguished scientists from several units. For further information concern- ing work in these areas, see the respective program descriptions in the Programs and Courses section of this catalog or contact the CNAS Graduate Student Affairs Center, at (800) 735-0717.